Nyctosaurus portrait in pencil. This magnificent animal lived during the late Cretaceous era, some 85 million years ago, near the American seashores. It belongs to the highly successful group of “modern Pterosaurs” with short tails and very long, untoothed beaks. It had a 2 meter wingspan and is considered to have been a soaring, gliding piscavore. I imagined it as having strong countershading much like a Sooty Tern. It is shown here preening.